Camberwell Beauty / Mourning Cloak

Jump to: Outline ssp. asopos ssp. borealis ssp. hyperborea ssp. lintnerii ssp. yedaluna

Outline

Including the Swedish Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Camberwell Beauty (Nymphalis antiopa) Camberwell BeautyNymphalis antiopa (Linnaeus, 1758)Mourning CloakNymphalis antiopa (Linnaeus, 1758) (see also Typusexemplar) described by Carl von Linné (1707-1778)Carl von Linné in 1758, the following subspecies and geographical races have been described:

Nymphalis antiopa ssp. antiopaNymphalis antiopa ssp. antiopa (Linnaeus, 1758)Nymphalis antiopa ssp. antiopa Linnaeus, 1758
Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos Fruhstorfer, 1909
Nymphalis antiopa ssp. borealisNymphalis antiopa ssp. borealis (Linnaeus, 1758)Nymphalis antiopa ssp. borealis Wnukowsky, 1927
Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea Seitz, 1914
Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii Fitch, 1856
Nymphalis antiopa ssp. yedalunaNymphalis antiopa ssp. yedaluna (Linnaeus, 1758)Nymphalis antiopa ssp. yedaluna Fruhstorfer, 1909

/PicturesNA/Graphics/Maps/antiopa_subspecies_map_world_medium.jpg

ssp. asopos

Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos Fruhstorfer, 1909

This subspecies occurs:

in JapanJapan
on Sakhalin, RussiaSakhalin
on the Kuril Islands, RussiaKuril Islands (north or north-east of JapanJapan)

It was described in 1909 by German Insect scientistEntomologist Hans Fruhstorfer (1866-1922)Hans Fruhstorfer in Fruhstorfer, 1909 - Neue Vanessa- und Pyrameis-Rassen:

Vanessa (Euvanessa) antiopa asopos nov. subspec.

The blue submarginal margins of the upper side of the wings are longer and narrower, the yellow distal margin somewhat more dusted than in Europeans, which also applies to the white and somewhat narrower outer margin on the underside.

Patria: Japan.

Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos and for comparison the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp.Nymphalis antiopa ssp. (Linnaeus, 1758)Nymphalis antiopa ssp.:

More specimens of the subspecies Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos see: Photos of pinned Camberwell beauties (Nymphalis antiopa): Nymphalis antiopa ssp. asopos

On the naming of the subspecies Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos see: Scientific Names

ssp. borealis

Nymphalis antiopa ssp. borealisNymphalis antiopa ssp. borealis (Linnaeus, 1758)Nymphalis antiopa ssp. borealis Wnukowsky, 1927

This subspecies occurs:

in RussiaRussia

It is smaller than the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp. antiopaNymphalis antiopa ssp. antiopa (Linnaeus, 1758)Nymphalis antiopa ssp. antiopa.

Woldemar WnukowskyWoldemar Wnukowsky describes the subspecies in Wnukowsky, 1927 - Neue Lepidopterenformen aus Nordost-Sibirien und dem Mongolischen Altai.:

Vanessa (Euvanessa) antiopa L. borealis subsp. nov.

Diagnosis: antiopa typ. similis, sed leviter minor (53-55 mm).

Patria: Siberia septentrionalis-orientalis (Jakutsk).

Typi in coli. Musei Zoolog. Universitatis Tomskensis (ad urbem Jakutsk, G. Antonovitsh leg.).

This north-east Siberian race differs from the typical representatives by its somewhat smaller size (53-55 mm). Described from specimens from the city of Yakutsk (G. Antonovich leg.). Types are in the collection of the Zoological Museum of Tomsk State University.

Nymphalis antiopa ssp. borealisNymphalis antiopa ssp. borealis (Linnaeus, 1758)Nymphalis antiopa ssp. borealis and for comparison the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp.Nymphalis antiopa ssp. (Linnaeus, 1758)Nymphalis antiopa ssp.:

More specimens of the subspecies Nymphalis antiopa ssp. borealisNymphalis antiopa ssp. borealis (Linnaeus, 1758)Nymphalis antiopa ssp. borealis see: Photos of pinned Camberwell beauties (Nymphalis antiopa): Nymphalis antiopa ssp. borealis

On the naming of the subspecies Nymphalis antiopa ssp. borealisNymphalis antiopa ssp. borealis (Linnaeus, 1758)Nymphalis antiopa ssp. borealis see: Scientific Names

ssp. hyperborea

Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea Seitz, 1914

This subspecies occurs:

in North America

It differs from the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp. antiopaNymphalis antiopa ssp. antiopa (Linnaeus, 1758)Nymphalis antiopa ssp. antiopa due to a smaller size, darker borders and larger spots Scott, 1986 - The Butterflies of North America - A Natural History and Field Guide.

Adalbert Seitz (1860-1938)Adalbert Seitz describes the subspecies in Seitz, 1924 - The Macrolepidoptera of the World: A systematic description of the hitherto known Macrolepidoptera:

...north, in Alaska. From there a specimen lies before me with very bright red-brown upper surface, white (not worn) margin slightly tinged with violet and on the under surface a light band across the middle of both wings; I figure it under the name hyperborea. rea form. nov. (93 f ).

Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea and for comparison the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp.Nymphalis antiopa ssp. (Linnaeus, 1758)Nymphalis antiopa ssp.:

Joseph BelicekJoseph Belicek analysed and described the subspecies Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea in more detail in 2013 Belicek, 2013 - Notes on the Holotype of Nymphalis antiopa hyperborea (Seitz, 1914). As a result, he significantly expanded the area of distribution usually cited until then (originally AlaskaAlaska Seitz, 1924 - The Macrolepidoptera of the World: A systematic description of the hitherto known Macrolepidoptera and northern Canada Canada Canada Canada):

...presently known from a large area on the North American continent. Ranging from Alaska, across Canada and the northern tier of states in the United States. Southward, specimens are known from North Carolina.

More specimens of the subspecies Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea see: Photos of pinned Camberwell beauties (Nymphalis antiopa): Nymphalis antiopa ssp. hyperborea

On the naming of the subspecies Nymphalis antiopa ssp. hyperboreaNymphalis antiopa ssp. hyperborea (Linnaeus, 1758)Nymphalis antiopa ssp. hyperborea see: Scientific Names

ssp. lintnerii

Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii Fitch, 1856

This subspecies occurs:

in North America

It was identified by I. A. LintnerI. A. Lintner in the US state of New York, /PicturesNA/Flags/us.png United StatesNew York Fitch, 1856 - Third Report on the Noxious, Beneficial and Other Insects of the State of New York. The wings are darker with a wider yellow border and smaller blue spots Rebel, 1910 - Fr. Berge's Schmetterlingsbuch nach dem gegenwärtigen Stande der Lepidopterologie. The subspecies was previously also partially categorised as A genetic or environmentally produced variation on the usual form of the speciesaberration (ab. lintnéri Fitch) Rebel, 1910 - Fr. Berge's Schmetterlingsbuch nach dem gegenwärtigen Stande der Lepidopterologie.

Asa Fitch (1809-1879)Asa Fitch writes in Fitch, 1856 - Third Report on the Noxious, Beneficial and Other Insects of the State of New York:

... This butterfly is closely related to tha Antiopa or White-bordered butterfly, a species which is common upon both sides of the Atlantic. Its wings have perfectly the same form and are similarly colored to those of the Antiopa, but their pale border is twice as broad as in that species, occupying a third of the length of the wings, and it is wholly destitute of the row of blue spots which occur in Antiopa forward of the border. Its ground color is deep rusty brown, much more tinged with liver-reddish than in Antiopa. ...

Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii and for comparison the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp.Nymphalis antiopa ssp. (Linnaeus, 1758)Nymphalis antiopa ssp.:

1887 Arthur Gardiner Butler (1844-1924)Arthur Gardiner Butler assigned Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii as A genetic or environmentally produced variation on the usual form of the speciesaberration, which he also found to be very similar to the A genetic or environmentally produced variation on the usual form of the speciesaberration Nymphalis antiopa f. hygiaeaNymphalis antiopa f. hygiaea (Linnaeus, 1758)Nymphalis antiopa f. hygiaea Butler, 1887 - Description of a new Butterfly allied to Vanessa antiopa:

The forms differing from type of V. antiopa - V. hygiaea, from Europe, and V.Lintneri, from the United States - are regarded, with every likelihood of being right, as aberrations of the species; judging from Hübner's figure (lettered "antiopa") and Fitch's description, thees aberrations appear to be very similar, differing chiefly from the normal for in the absence of many of th blue spots and the slightly wider yellow border of all the wings.

More specimens of the subspecies Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii see: Photos of pinned Camberwell beauties (Nymphalis antiopa): Nymphalis antiopa ssp. lintnerii

On the naming of the subspecies Nymphalis antiopa ssp. lintneriiNymphalis antiopa ssp. lintnerii (Linnaeus, 1758)Nymphalis antiopa ssp. lintnerii see: Scientific Names

ssp. yedaluna

Nymphalis antiopa ssp. yedalunaNymphalis antiopa ssp. yedaluna (Linnaeus, 1758)Nymphalis antiopa ssp. yedaluna Fruhstorfer, 1909

This subspecies occurs (Quelle: ENVIS (Environmental Information System) on Faunal Diversity (2011)):

in Chumbi-Valley, Tibet, ChinaChumbi-Valley
in Triangle IndiaIndia/Tibet, ChinaTibet/BhutanBhutan
in Manipur, IndiaManipur

It was described in 1909 by the German entomologist Hans Fruhstorfer (1866-1922)Hans Fruhstorfer and caught by him at the Chumbi Pass in July/August 1912. It is a subspecies with only slight variation, the occurrence of which has declined significantly.

Hans Fruhstorfer (1866-1922)Hans Fruhstorfer describes the subspecies in Fruhstorfer, 1909 - Neue Vanessa- und Pyrameis-Rassen:

Vanessa antiopa yedanula nov. subspec.

(Yedanula Name of the deceased Empress of China.)

Smaller than the previous one Nymphalis antiopa ssp. asoposNymphalis antiopa ssp. asopos (Linnaeus, 1758)Nymphalis antiopa ssp. asopos; the white outer border on the underside is narrower, even more densely dotted in black; Distal margin of the upper side darker yellow than in Europeans; the subapical spots are narrower, more reminiscent of the American form.

Patria: China, Szetchuan, Type of Siao-Lou. Of Europeans, South Tyroleans ♂ ♂ are the largest, those from Finland and the Urals the smallest.

Nymphalis antiopa ssp. yedalunaNymphalis antiopa ssp. yedaluna (Linnaeus, 1758)Nymphalis antiopa ssp. yedaluna and for comparison the Nominiform or (new) nominotypic taxon: Taxon defined by the same name-bearing type as the higher-ranking taxon to which it itself belongs.Type Species Nymphalis antiopa ssp.Nymphalis antiopa ssp. (Linnaeus, 1758)Nymphalis antiopa ssp.:

More specimens of the subspecies Nymphalis antiopa ssp. yedalunaNymphalis antiopa ssp. yedaluna (Linnaeus, 1758)Nymphalis antiopa ssp. yedaluna see: Photos of pinned Camberwell beauties (Nymphalis antiopa): Nymphalis antiopa ssp. yedaluna

On the naming of the subspecies Nymphalis antiopa ssp. yedalunaNymphalis antiopa ssp. yedaluna (Linnaeus, 1758)Nymphalis antiopa ssp. yedaluna see: Scientific Names

end of February: Camberwell Beauty (Nymphalis antiopa)
GermanyGermany		rare, hibernating
Brandenburg		common, hibernating
Berlin		common, hibernating
Bremen		extinct
Baden-Württemberg		common, hibernating
Bavaria		common, hibernating
Hesse		very rare, hibernating
Hamburg		common, hibernating
Mecklenburg-Vorpommern		common, hibernating
Lower Saxony		common, hibernating
North Rhine-Westphalia		very rare, hibernating
Rhineland-Palatinate		rare, hibernating
Schleswig-Holstein		rare, hibernating
Saarland		common, hibernating
Saxony		common, hibernating
Saxony Anhalt		common, hibernating
Thuringia		common, hibernating